The Queen has held talks with US President Donald Trump ahead of American Independence Day.
The two heads of state chatted via telephone, just over six months since their last meeting at a Buckingham Palace reception marking the 70th anniversary of NATO.
"The president wished the Queen a happy birthday, marking 94 extraordinary years," the White House said in a statement.
"The president also expressed his condolences for the British people who have lost their lives during the coronavirus pandemic.
"The president and the Queen discussed close co-operation on defeating the virus and reopening global economies.
"The president and the Queen also reaffirmed that the United States and United Kingdom stand together in our special relationship and will emerge from this trying time stronger than ever before."
A tweet posted on the official royal family account on Tuesday also confirmed the call.
"Today, The Queen spoke to President Trump by telephone from Windsor Castle ahead of Independence Day in the United States on the 4th July," the post said.
Following Trump's three-day state visit in June 2019 he described the Queen as a "spectacular woman" in an interview with a US cable network and claimed they had "chemistry" .
"The meeting with the Queen was incredible. I think I can say I've really got to know her because I've sat with her many times and we had automatic chemistry," he told Fox News.
"You understand that feeling, it's a good feeling. But she's a spectacular woman".
The Queen has held telephone conversations with a series of leaders during the coronavirus lockdown, including French President Emmanuel Macron, Australian Prime Minister Scott Morrison and Justin Trudeau, Canada's prime minister.
Trump's conversation with the Queen came ahead of his country's Independence Day celebrations on July 4.
A White House event marking the anniversary is being held despite concerns expressed by some US politicians about the number of people expected to attend during the coronavirus outbreak.
World leaders, including Trump, were hosted by the Queen at the NATO reception staged at her official London residence in December.
Earlier last year, the US president was welcomed on a three-day state visit by the Queen during a ceremony at Buckingham Palace on June 3.